Summer Short Course

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Summer Short Courses in Organismic Plant Biology

The Leaf

June 5 – 12, 2019

The 2019 short course in organismic plant biology at the Arnold Arboretum of Harvard University will bring together a group of instructors to lead a broadly integrative analysis of the structure and function of leaves. Topics to be covered will include organogenesis and morphogenesis, evolutionary history of leaves, leaf anatomy, leaf traits, ecophysiology, physiology, hydraulics, stomatal functioning, and gas exchange. This course will alternate between lectures, laboratory work, and opportunities to examine state-of-the art methodologies. By the end of the course, students should expect to have acquired a foundational knowledge of how leaves develop and interact with the environment.

Summer Short Courses are free for all participants.

Download a poster/flyer (feel free to print and distribute!)

Course Information


Craig Brodersen (Yale University)
Pamela Diggle (University of Connecticut)
William (Ned) Friedman (Harvard University)
Noel M. Holbrook (Harvard University)
Cynthia Jones (University of Connecticut)
Fulton Rockwell (Harvard University)


Summer Short Courses are open to postdoctoral researchers, graduate students, and undergraduates in their final year of study (who have been admitted to a graduate or professional program for the fall of 2019).

Non-US-citizens are welcome to apply. Please note that if you are not a United States citizen, you are responsible for obtaining the appropriate visa to travel to the United States to attend the course.

Travel Assistance and Lodging for Participants

Lodging and week-day meals will be provided for all course participants. All participants are eligible to receive up to $500 (domestic travel) or $700 (international travel) USD of funds to help pay for travel to/from the course.

Application Information

Application Deadline

Applications must be submitted by 11:30 pm March 15th, 2019.

How to Apply

Apply online by clicking here.

You will have a register an account in order to apply. There are a few steps you will need to complete:

1) Please click the Submission link and follow the instructions to register for an account.

2) Search for grant listings for the “Summer Short Course – The Leaf” .

3) Click “Apply” or “Add to Cart”. (if you use “Add to Cart”, the listing will now show up when you click on the “My Interest Cart” tab).

4) Follow application instructions. The application includes a way for you to send referee letter requests (submission instructions will be sent to your referees). Be sure to click “Final Review and Submit” when you are ready to submit!

Required Application Materials

1) Contact information

2) Academic information

3) Essay: In a total of 1000 words or fewer, please tell us:
A) Why you want to participate in the course?
B) What are your scientific interests?
C) While no experience is required, please describe your background in plant biology.

4) Contact information (name and email address) for two referees. As part of the online submission, you will be asked to send a request to the referees to submit a letter of recommendation. Each referee will be automatically emailed with a link to an online submission form where they can upload a letter of recommendation. Both referees must upload their letter of recommendation via the link by March 15th at 11:30 pm.

Application Evaluation

Course instructors are charged with evaluating applications. We anticipate announcing application status by mid-April.

Additional Information

Summer Short Courses in organismic plant biology are sponsored by the Arnold Arboretum of Harvard University, the Department of Organismic and Evolutionary Biology, and New Phytologist Trust.


Please contact Pam Diggle for any questions.

Submit Application